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By screening in four different media conditions, we uncovered their metabolic regulation, including the fitness defects in the mitochondrial folate transporter SLC25A32 KO cells that can be buffered in galactose condition due to substrate limitation in de novo purine biosynthesis

20 The folate and methionine cycles are interlinked by the rate-limiting enzyme methionine synthase (MTR), and are essential to produce SAM, the universal methyl donor (Figure 3A)