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As research on transcription and translation advanced, it was discovered that initial RNA transcripts typically require intron removal and the linkage of exons to form mature messenger RNA (mRNA), a process known as RNA splicing

In vitro studies have shown that elevating cytoplasmic Ca 2+ with the calcium ionophore A23187 led to a 2.9-fold upregulation of the nuclear-encoded NDUFA9, mitochondrial COX protein, and mitochondrial transcription factor TFAM [239, 240]

Notably, these observations were found in a relatively short time after Gclc deletion (i.e., three weeks)

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